[1] 黄金霞,王亮生,李晓梅,等.花色变异的分子基础与进化模式研究进展[J].植物学通报,2006, 23(4): 321-333.
[2] Holton T A, Cornish E C. Genetics and biochemistry of anthocyanin biosynthesis[J]. Plant Cell, 1995, 7(7): 1071-1083.
[3] Ramsay N A, Glover B J. MYB-bHLH-WD40 protein complex and the evolution of cellular diversity[J]. Trends Plant Sci, 2005,10(2): 63-70.
[4] 张全琪,朱家红,倪燕妹,等.植物bHLH转录因子的结构特点及其生物学功能[J].热带亚热带植物学报, 2000,19(1):84-90.
[5] Nesi N, Debeaujon I, Jond C, et al. The TT8 gene encodes a basic helix-loop-helix domain protein required for expression of DFR and BAN genes in Arabidopsis siliques[J]. Plant Cell, 2000, 12 (10): 1863-1878.
[6] Heisler MG, Atkinson A, Bylstra YH, et al. SPATULA, a gene that controls development of carpel margin tissues in Arabidopsis, encodes a bHLH protein[J]. Development,2001,128:1089-1098.
[7] Sorensen A M, Kröber S, Unte U S, et al. The Arabidopsis ABORTED MICROSPORES (AMS) gene encodes a MYC class transcription factor[J]. Plant J,2003, 33(2): 413-423.
[8] Leivar P, Monte E, Oka Y, et al. Multiple phytochrome-interacting bHLH transcription factors repress premature seedling photomorphogenesis in darkness[J]. Curr Biol,2008, 18 (23): 1815-1823.
[9] Abe H, Urao T, Ito T, et al. Arabidopsis AtMYC2 (bHLH) and AtMYB2 (MYB) function as transcriptional activators in abscisic acid signaling[J]. Plant Cell, 2003,15 (1): 63-78.
[10] Lorenzo O, Chico JM, Sánchez-Serrano JJ, et al. JASMONATE-INSENSITIVE1 encodes a MYC transcription factor essential to discriminate between different jasmonate-regulated defense responses in Arabidopsis[J]. Plant Cell, 2004,16 (7): 1938-1950.
[11] Li H, Sun J, Xu Y, et al. The bHLH-type transcription factor AtAIB positively regulates ABA response in Arabidopsis. Plant Mol Biol.2007, 65 (5): 655-665.
[12] Bou-Torrent J, Roig-Villanova I, Galstyan A, et al. PAR1 and PAR2 integrate shade and hormone transcriptional networks[J]. Plant Signal Behav. 2008, 3 (7): 453-454.
[13] Rampey R A, Woodward AW, et al. An Arabidopsis basic helix-loop-helix leucine zipper protein modulates metal homeostasis and auxin conjugate responsiveness[J]. Genetics. 2006,174 (4): 1841-1857.
[14] Long TA, Tsukagoshi H, Busch W, et al. The bHLH transcription factor POPEYE regulates response to iron deficiency in Arabidopsis roots[J]. Plant Cell,2010,22 (7): 2219-2236.
[15] Ludwig SR, Habera LF, Dellaporta SL, et al. Lc, a member of the maize R gene family responsible for tissuespecific anthocyanin production, encodes a protein similar to transcriptional activators and contains the myc-homology region[J]. Proc Natl Acad Sci USA, 1998,86 (18): 7092-7096.
[16] ChandlerV L, Radieella J P, Robbins T P, et al. Two regulatory genes of the maize anthocyanin pathway are homologous:Isolation of B utilizing R genomic sequenees[J]. Plant Cell,1989,1(12):1175-1183.
[17] Goff S A, Cone K C, Chandler V L. Functional analysis of the transcriptional activator encoded by the maize B gene: evidence for a direct functional interaction between two classes of regulatory proteins[J]. Genes Dev., 1992,6 (5): 864-875.
[18] Goodrich J, Carpenter R and Coen E S. 1992. A common gene regulates pigmentation patten in diverse plant species. Cell, 68(5):955-964.
[19] Elomaa P, Mehto M, Kotilainen M, et al. A bHLH transcription factor mediates organ, region and flower type specific signals on dihydrofl avonol-4-reductase (dfr) gene expression in the inflorescence of Gerbera hybrida (Asteraceae). Plant J., 1998,16(1):93-99.
[20] Sakamoto W, Ohmori T, Kageyama K,et al. The purple leaf (Pl) locus of rice:The pl allele has a complex organization and includes two Genes encoding basie helix-loop-helix proteins involved in anthocyanin biosynthesis. Plant Cell Physiol, 2001, 42(9):982-991.
[21] Park K, Ishikawa N, Morita Y, et al. A bHLH regulatory gene in the common morning glory, Ipomoea Purpurea, controls anthocyanin biosynthesis in flowers, proanthocyanidin and phytolnelanin pigmentation in seeds, and seed trichome formation.The Plant Joumal, 2007b, 49(4):641-654
[22] Nakatsuka T, Haruta K S, Pitaksutheepong C, et al. Identification and characterization of R2R3-MYB and bHLH transcription factors regulating anthocyanin biosynthesis in gentian flowers[J]. Plant Cell Physiol. 2008, 49: 1818-1829.
[23] Espley R V, Hellens R P, Putterill J, et al. Red colouration in apple fruit due to the activity of the MYB transcription factor, MdMYB10[J]. Plant J.,2007,49:414-427.
[24] Franken P., Schrell S., Peterson P A., et al. Molecular analysis of protein domain function encoded by the myb-homologous maize genes CI, Zm1 and Zm 38[J]. The Plant Journal,1994,6 (1):21-30.
[25] Spelt C, Quattrocchio F, Mol JNM, et al.Anthocyanin1 of petunia encodes a basic helix-loop-helix protein that directly activates transcription of structural anthocyanin genes[J]. Plant Cell, 2000,12 (9): 1619-1631.
[26] Yamagishi M, Shimoyamada Y, Nakatsuka T, et al. Two R2R3-MYB genes, homologs of petunia AN2, regulate anthocyanin biosyntheses in flower tepals, tepal spots and leaves of Asiatic hybrid lily[J]. Plant Cell Physiol, 2010, 51:463-474.
[27] Wang Kui-lin, Bolitho, K., Grafton, K., et al. An R2R3 MYB transcription factor associated with regulation of the anthocyanin biosynthetic pathway in Rosaceae[J]. BMC Plant Biology, 2010,10: 50.
[28] 王 勇,陈克平,姚 勤. bHLH转录因子家族研究进展[J].遗传,2008, 30 (7): 821-830.
[29] 方文培. 中国芍药属植物的研究[J]. 植物分类学报, 1958, 7(4): 297-323.
[30] 王莲英. 中国牡丹品种图志[M]. 北京: 中国林业出版社, 1997, 2-7.
[31] 李嘉珏. 中国牡丹与芍药[M]. 北京: 中国林业出版社, 1999.
[32] 龚 询.滇牡丹复合群体分类研究[J]. 7 昆明植物研究所,1990.
[33] 王志芳, 王 雁, 岳 桦. 珍稀资源: 黄牡丹[J].中国城市林业,2007, 5(2): 59- 60.
[34] Zhou L, Wang Y, Ren L, et al. Overexpression of Pl-CHI1, a homologue of the chalcone isomerase gene from tree peony (Paeonia suffruticosa), reduces the intensity of flower pigmentation in transgenic tobacco[J]. Plant Cell, Tissue and Organ Culture, 2014, 16 (3): 285-295.
[35] Zhou L, Wang Y, Peng Z. Molecular characterization and expression analysis of chalcone synthase gene during flower development in tree peony (Paeonia suffruticosa)[J]. Afr J Biotechnol, 2011, 10(8):1275-1284.
[36] Fan C, Purugganan M D, Thomas D T, et al. Heterogeneous evolution of the Myc-like Anthocyanin regulatory gene and its phylogenetic utility in Cornus L.(Cornaceae)[J]. Mol Phylogenet Evol,2004, 33(3): 580-594.
[37] Jiang Y, Deyholos M K. Comprehensive transcriptional profiling of NaCl-stressed Arabidopsis roots reveals novel classes of responsive genes[J]. BMC Plant Biol, 2006, 6(1): 25.
[38] Jiang Y, Yang B, Deyholos M K. Functional characterization of the Arabidopsis bHLH92 transcription factor in abiotic stress[J]. Mol Genet Genomics, 2009, 282(5): 503-516.
[39] Li XX,Duan XP,Jiang HX, et al. Genome-wide analysis of basic/helix-loop-helix transcription factor family in rice and Arabidopsis[J]. Plant Physiology, 2006, 141(4):1167-1184.
[40] Hichri I, Heppel S C, Pillet J, et al. The basic helix-loop-helix transcription factor MYC1 is involved in the regulation of the flavonoid biosynthesis pathway in grapevine[J]. Mol Plant, 2010, 3(3): 509-523.
[41] Xie X B, Li S, Zhang R F, et al. The bHLH transcription factor MdbHLH3 promotes anthocyanin accumulation and fruit colouration in response to low temperature in apples[J]. Plant Cell Environ, 2012, 35(11): 1884-1897.
[42] Quattrocchio F, Wing J F, Va K, et al. Analysis of bHLH and MYB domain proteins: species-specific regulatory differences are caused by divergent evolution of target anthocyanin genes[J]. Plant J, 1998,13(4): 475-488.
[43] Martin C, Prescott A, Mackay S, et al. Control of anthocyanin biosynthesis in flowers of Antirrhinum majus. Plant J.1991,1 (1): 37-49.
[44] Quattrocchio F,Wing J F,Leppen H,et al. Regulatory genes controlling anthocyanin pigmentation are functionally conserved among plant species and have distinct sets of target genes[J]. The Plant Cell, 1993, 5 (11): 1497-1512.
[45] Quattrocchio F, Baudry A, Lepiniec L, et al. The regulation of flavonoid biosynthesis. In: Grotewold E(ed) The science of flavonoids[J]. New York, Springer, pp.2006: 97-122.
[46] de Vetten N, Quattrocchio F, Mol J, et al. The an11 locus controlling flower pigmentation in petunia encodes a novel WD-repeat protein conserved in yeast, plants, and animals[J]. Genes & Development, 1997,11:1422-1434.
[47] Elomaa P, Mehto M, Kotilainen M, et al. A bHLH transcription factor mediates organ, region and flower type specific signals on dihydrofl avonol-4-reductase (dfr) gene expression in the inflorescence of Gerbera hybrida (Asteraceae). Plant J., 1998,16(1):93-99.
[48] Li C, Du H, Wang L, et al. Flavonoid composition and antioxidant activity of tree peony (Paeonia section Moutan) yellow flowers[J]. J Agric Food Chem, 2009, 57: 8496-8503.
[49] 周 琳,王 雁, 律春燕,等.云南野生黄牡丹花色素成分的鉴定[J].东北林业大学学报,2011, 39(8):52-54.
[50] Wang LS, Hashimoto F, Shiraishi A, et al. Phenetics in tree peony species from China by flower pigment cluster analysis[J].J Plant Res, 2001a, 114: 213-221.
[51] Wang LS, Shiraishi A, Hashimoto F, et al. Analysis of petal anthocyanins to investigate flower coloration of Zhongyuan (Chinese) and Daikon Island (Japanese) tree peony cultivars[J]. J Plant Res,2001b, 114:33-43.
[52] 韩科厅,赵 莉,唐杏姣, 等.菊花花色素苷合成关键基因表达与花色表型的关系[J].园艺学报, 2012,39 (3):516-524.