Spatial Distribution Patterns and Associations of Dominate Population 
in Two Types of Broadleaved Korean Pine Mixed Forest

ZHAO Zhong-hua; HUI Gang-ying; HU Yan-bo; ZHOU Hong-min; ZHANG Lian-jin

Volume 24 Issue 5

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Spatial Distribution Patterns and Associations of Dominate Population in Two Types of Broadleaved Korean Pine Mixed Forest

1.
Research Institute of Forestry, Chinese Academy of Forestry, Key Laboratory of Tree Breeding and Cultivation, State Forestry Administration, Beijing 100091, China
2.
Longquan Research Institute of Forestry, Longquan 323700, Zhejiang, China

Received Date: 2011-02-12

Abstract

Based on the location data for each timber from two broadleaved Korean pine (Pinus koraiensis) mixed forest plots which area is 1 hm2 at Jiaohe of Jilin province, the spatial distribution pattern and tree species spatial correlation of P. koraiensis and main dominate population in middle and upper vertical forest layer were chosen as study objects, the O-ring statistics based on RipleyK(d) function and Mark correlation function was used to analyze the spatial distribution and spatial correlation. The result showed that spatial distribution pattern of P. koraiensis and Fraxinus mandshurica were clumped at smaller scales in F. mandshurica-P. koraiensis forest, P. koraiensis was especially in middle vertical layer. P. koraiensis and Tilia amurensis spatial distribution pattern were clumped at multiple scales. At same vertical layer, P. koraiensis had no significant spatial association to F. mandshurica and T. amurensis, however, there had some association in different vertical layer of different population. P. koraiensis in upper layer has significant negative correlation with midstory layer at 0～19 m, this properties was not obviously in T. amurensis-P. koraiensis mixed forest. F. mandshurica and T. amurensis in upper vertical layer showed significant negative correlation with midstory layer at smaller scales. P. koraiensis in upper layer shower significant negative correlation with F. mandshurica and T. amurensis in middle vertical layer lager scale, but P. koraiensis in middle vertical layer had no correlation with F. mandshurica and T. amurensis in upper layer.

References

[1]	宋永昌. 植被生态学[M].上海:华东师范大学出版社,2001
[2]	张金屯,孟东平. 芦芽山华北落叶松林不同龄级立木的点格局分析[J].生态学报,2004,24(1): 35-40
[3]	张金屯. 群落二维格局分析的两种方法[J]. 西北植物学报,2004, 24(8): 1448-1451
[4]	张会儒,汤孟平,舒清态.森林生态采伐的理论与实践[M].北京:中国林业出版社, 2006
[5]	叶芳,彭世揆.种群空间分布理论的发展历史及其现状[J].林业资源管理, 1997(6): 55-58
[6]	惠刚盈,李丽,赵中华,等.林木空间分布格局研究方法[J].生态学报,2007,27(11):4717-4728
[7]	谢宗强,陈伟烈,刘正宇.银杉种群的空间分布格局[J].植物学报, 1999, 41(1): 95-101
[8]	王太鑫,丁雨龙,刘永建.巴西木竹无性系种群的分布格局[J].南京林业大学学报:自然科学版,2005,29(3): 37-40
[9]	张金屯. 植物种群空间分布的点格局分析[J].植物生态学报,1998, 22(4): 344-349
[10]	Ripley B D. Spatial Statistics[M]. NewYork: Wiley,1981
[11]	Ripley B D.The second-order analysis of stationary point processes[J].Journal of Applied Probability, 1976,13:255-266
[12]	Ripley B D. Modelling spatial pattern[J]. Journal of the Royal Statistical Society. Series B, 1977,39: 172-212
[13]	汤孟平,唐守正,雷相东.Ripley’sK(d)函数分析种群空间分布格局的边缘校正[J].生态学报, 2003,23(8): 1533-1538
[14]	惠刚盈.角尺度——一个描述林木个体分布格局的结构参数[J].林业科学,1999,35(1): 37-42
[15]	徐海,惠刚盈,胡艳波,等.天然红松阔叶林不同径阶林木的空间分布特征分析[J].林业科学研究, 2006,19(6): 687-691
[16]	王蕾,张春雨,赵秀海.长白山阔叶红松林的空间分布格局[J].林业科学,2009,45(5):55-60
[17]	侯向阳,韩进轩.长白山红松林主要树种空间格局的模拟分析[J].植物生态学报,1997, 21(3): 242-249
[18]	Wiegand T, Moloney K A. Rings, circles and null models for point pattern analysis in ecology[J].Oikos, 2004,104: 209-229
[19]	徐化成.中国红松阔叶林[M].北京:中国林业出版社,2001
[20]	阳含熙.长白山北坡红松阔叶林主要树种的分布格局[M].森林生态系统研究, 1985(1): 1-14
[21]	张健,郝占庆,宋波,等. 长白山阔叶红松林中红松与紫椴的空间分布格局及其关联性[J]. 应用生态学报,2007,18(8):1681-1687
[22]	覃林.统计生态学[M],北京:中国林业出版社,2009
[23]	汤孟平,周国模,施拥军,等.天目山常绿阔叶林优势种群及其空间分布格局[J]. 植物生态学报,2006,30(5):743-752
[24]	Condit R,Ashton P S,Baker P, et al. Spatial patterns in the distribution of tropical tree species[J].Science, 2000,288:1414-1418
[25]	Schurr F M,Bossdorf O,Milton S J, et al.Spatial pattern formation in semi-arid shrub land: a priori predicted versus observed pattern characteristics[J]. Plant Ecology, 2004,173:271-282
[26]	喻泓,杨晓晖,慈龙骏.地表火对红花尔基沙地樟子松种群空间分布格局的影响[J]. 植物生态学报,2009,33(1):71-80
[27]	缪宁,史作民,冯秋红,等. 川西亚高山岷江冷杉种群的空间格局分析[J].林业科学,2008,44(12):1-5
[28]	Yu H, Wiegand T, Yang X, et al. The impact of fire and density-dependent mortality on the spatial patterns of a pine forest in the Hulun Buir sandland, Inner Mongolia, China[J]. Forest Ecology and Management, 2009, 257(10): 2098-2107
[29]	臧润国.红松阔叶林林冠空隙动态的研究 .北京:北京林业大学,1995
[30]	臧润国,郭忠凌,高文韬.长白山自然保护区阔叶红松林林隙更新的研究[J].应用生态学报,1998,9(4):349-353
[31]	宋新章,张慧玲,肖文发,等. 长白山区阔叶红松林采伐林隙种子库研究[J].北京林业大学学报,2009,32(2):17-24
[32]	Nathan R. Long-distance dispersal of plants[J].Science, 2006, 313: 786-788
[33]	He F, Legendre P, La Frankie J V. Distribution patterns of tree species in a Malaysian tropical rain forest[J].Journal of Vegetation Science, 1997,8: 105-11

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通讯作者: 陈斌, bchen63@163.com

1.
沈阳化工大学材料科学与工程学院沈阳 110142

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Spatial Distribution Patterns and Associations of Dominate Population in Two Types of Broadleaved Korean Pine Mixed Forest

1. Research Institute of Forestry, Chinese Academy of Forestry, Key Laboratory of Tree Breeding and Cultivation, State Forestry Administration, Beijing 100091, China

2. Longquan Research Institute of Forestry, Longquan 323700, Zhejiang, China

Received Date: 2011-02-12

Abstract: Based on the location data for each timber from two broadleaved Korean pine (Pinus koraiensis) mixed forest plots which area is 1 hm2 at Jiaohe of Jilin province, the spatial distribution pattern and tree species spatial correlation of P. koraiensis and main dominate population in middle and upper vertical forest layer were chosen as study objects, the O-ring statistics based on RipleyK(d) function and Mark correlation function was used to analyze the spatial distribution and spatial correlation. The result showed that spatial distribution pattern of P. koraiensis and Fraxinus mandshurica were clumped at smaller scales in F. mandshurica-P. koraiensis forest, P. koraiensis was especially in middle vertical layer. P. koraiensis and Tilia amurensis spatial distribution pattern were clumped at multiple scales. At same vertical layer, P. koraiensis had no significant spatial association to F. mandshurica and T. amurensis, however, there had some association in different vertical layer of different population. P. koraiensis in upper layer has significant negative correlation with midstory layer at 0～19 m, this properties was not obviously in T. amurensis-P. koraiensis mixed forest. F. mandshurica and T. amurensis in upper vertical layer showed significant negative correlation with midstory layer at smaller scales. P. koraiensis in upper layer shower significant negative correlation with F. mandshurica and T. amurensis in middle vertical layer lager scale, but P. koraiensis in middle vertical layer had no correlation with F. mandshurica and T. amurensis in upper layer.

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Reference (33)

[1]	宋永昌. 植被生态学[M].上海:华东师范大学出版社,2001
[2]	张金屯,孟东平. 芦芽山华北落叶松林不同龄级立木的点格局分析[J].生态学报,2004,24(1): 35-40
[3]	张金屯. 群落二维格局分析的两种方法[J]. 西北植物学报,2004, 24(8): 1448-1451
[4]	张会儒,汤孟平,舒清态.森林生态采伐的理论与实践[M].北京:中国林业出版社, 2006
[5]	叶芳,彭世揆.种群空间分布理论的发展历史及其现状[J].林业资源管理, 1997(6): 55-58
[6]	惠刚盈,李丽,赵中华,等.林木空间分布格局研究方法[J].生态学报,2007,27(11):4717-4728
[7]	谢宗强,陈伟烈,刘正宇.银杉种群的空间分布格局[J].植物学报, 1999, 41(1): 95-101
[8]	王太鑫,丁雨龙,刘永建.巴西木竹无性系种群的分布格局[J].南京林业大学学报:自然科学版,2005,29(3): 37-40
[9]	张金屯. 植物种群空间分布的点格局分析[J].植物生态学报,1998, 22(4): 344-349
[10]	Ripley B D. Spatial Statistics[M]. NewYork: Wiley,1981
[11]	Ripley B D.The second-order analysis of stationary point processes[J].Journal of Applied Probability, 1976,13:255-266
[12]	Ripley B D. Modelling spatial pattern[J]. Journal of the Royal Statistical Society. Series B, 1977,39: 172-212
[13]	汤孟平,唐守正,雷相东.Ripley’sK(d)函数分析种群空间分布格局的边缘校正[J].生态学报, 2003,23(8): 1533-1538
[14]	惠刚盈.角尺度——一个描述林木个体分布格局的结构参数[J].林业科学,1999,35(1): 37-42
[15]	徐海,惠刚盈,胡艳波,等.天然红松阔叶林不同径阶林木的空间分布特征分析[J].林业科学研究, 2006,19(6): 687-691
[16]	王蕾,张春雨,赵秀海.长白山阔叶红松林的空间分布格局[J].林业科学,2009,45(5):55-60
[17]	侯向阳,韩进轩.长白山红松林主要树种空间格局的模拟分析[J].植物生态学报,1997, 21(3): 242-249
[18]	Wiegand T, Moloney K A. Rings, circles and null models for point pattern analysis in ecology[J].Oikos, 2004,104: 209-229
[19]	徐化成.中国红松阔叶林[M].北京:中国林业出版社,2001
[20]	阳含熙.长白山北坡红松阔叶林主要树种的分布格局[M].森林生态系统研究, 1985(1): 1-14
[21]	张健,郝占庆,宋波,等. 长白山阔叶红松林中红松与紫椴的空间分布格局及其关联性[J]. 应用生态学报,2007,18(8):1681-1687
[22]	覃林.统计生态学[M],北京:中国林业出版社,2009
[23]	汤孟平,周国模,施拥军,等.天目山常绿阔叶林优势种群及其空间分布格局[J]. 植物生态学报,2006,30(5):743-752
[24]	Condit R,Ashton P S,Baker P, et al. Spatial patterns in the distribution of tropical tree species[J].Science, 2000,288:1414-1418
[25]	Schurr F M,Bossdorf O,Milton S J, et al.Spatial pattern formation in semi-arid shrub land: a priori predicted versus observed pattern characteristics[J]. Plant Ecology, 2004,173:271-282
[26]	喻泓,杨晓晖,慈龙骏.地表火对红花尔基沙地樟子松种群空间分布格局的影响[J]. 植物生态学报,2009,33(1):71-80
[27]	缪宁,史作民,冯秋红,等. 川西亚高山岷江冷杉种群的空间格局分析[J].林业科学,2008,44(12):1-5
[28]	Yu H, Wiegand T, Yang X, et al. The impact of fire and density-dependent mortality on the spatial patterns of a pine forest in the Hulun Buir sandland, Inner Mongolia, China[J]. Forest Ecology and Management, 2009, 257(10): 2098-2107
[29]	臧润国.红松阔叶林林冠空隙动态的研究 .北京:北京林业大学,1995
[30]	臧润国,郭忠凌,高文韬.长白山自然保护区阔叶红松林林隙更新的研究[J].应用生态学报,1998,9(4):349-353
[31]	宋新章,张慧玲,肖文发,等. 长白山区阔叶红松林采伐林隙种子库研究[J].北京林业大学学报,2009,32(2):17-24
[32]	Nathan R. Long-distance dispersal of plants[J].Science, 2006, 313: 786-788
[33]	He F, Legendre P, La Frankie J V. Distribution patterns of tree species in a Malaysian tropical rain forest[J].Journal of Vegetation Science, 1997,8: 105-11

Spatial Distribution Patterns and Associations of Dominate Population in Two Types of Broadleaved Korean Pine Mixed Forest

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Spatial Distribution Patterns and Associations of Dominate Population in Two Types of Broadleaved Korean Pine Mixed Forest

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